Here is information about my academic life. For blog posts with advice for undergraduates or students currently pursuing Masters or PhD’s, check out the Blog page. My Google Scholar profile can be found here.


  1. Twomey CR, Hartnett AT, Sosna MMG, Romanczuk P. 2020. Searching for structure in collective systems. Theory in Biosciences.

  2. Sosna MMG, Twomey CR, Bak-Coleman J, Poel W, Daniels BC, Romanczuk P, Couzin ID. 2019. Individual and collective encoding of risk in animal groups. Proceedings of the National Academy of Sciences. 116(41): 20556-20561
  3. Stuber EF, Grobis MM, Abbey-Lee R, Kempenaers B, Mueller JC, Dingemanse NJ. 2014. Perceived predation risk affects sleep behaviour in great tits (Parus major). Animal Behaviour. 98: 157-165.

  4. Grobis MM, Pearish SP, Bell AM. 2013. Avoidance or escape? Discriminating between two hypotheses for the function of schooling in threespine sticklebacks. Animal Behaviour. 85(1): 187-194.

In progress

  1. Davidson JD*, Sosna MMG*, Twomey CR, Sridhar VH, Couzin ID. Collective detection based on visual information in fish schools. in prep.

  2. Jolles JW*, Sosna MMG*, Jolles JW, Mazué GPF, Twomey CR, Bak-Coleman J, Rubenstein DI, Couzin ID. Quantifying the spatial distribution of risk in animal groups. in prep.

  3. Twomey CR, Sosna MMG, Ragazzo G, Bak-Coleman J, Couzin ID. Between-group dynamics and emergent social forces at the limits of sensory perception. in prep.

* denotes joint first authorship